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A portion of it is done by the American cirno poster, who necrobumps threads with short replies when the proxyfag isn't active. He also has the same spam thread creation method (copy-pasting threads from /a/ or using the same few images that the proxyfag also uses).
Deduction: 1. The proxy spammer is coordinating with the cirnotroon meaning that there is a mafia controlling the board from the shadows
or 2. The proxy spammer is the cirnotroon and he uses the short reply method when he doesn't have access to his proxies and doesn't want to get his american ips banned.