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Marowak roam the deserts alone. Once every few years a male and female will meet, and they'll have a single child.
The male will then continue on his way, and the mother will take the child with her, raising it and protecting it.
At a certain point in the child's life the mother challenges it to a fight to the death. If it loses, the mother carries on until she finds another male. If the child wins, however, it must wear her skull until it is time to evolve, at which time that skull will bond to it.
That one from RBY wasn't special, it was just a crying little bitch.