>>56544481At first they were unrelated, they were representations of Japanese Kamis; Thunder, Wind, and Harvest.
It was not until they were given the form based on Chinese celestial beasts that they acquired the qualities of seasons, as well as cardinal points due to the different types of winds in each season, similar to the Greek Anemoi.
Thus:
Enamorus represents spring, mating season
Thundurus represents summer, typhoon season
Landorus represents autumn, harvest season
Tornadus represents winter, icy wind season