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The Tapus are Mimikyu which unnaturally evolved when they put on the mask relics of actual old guardian spirits. Similar to Rotom changing forms based on appliances, they lost their ghost typing and gained the type of the diety.
They were Mimikyu who tried to become beloved by donning such cherished items, unbeknownst to the Alolan People.
The claw seen of Mimikyu shares sharp resemblance to Fini's.
The realization that their gods are a species that are "mistreated" by most of the population is the source of the shock of the scholar who saw a Mimikyu's true form.
Tapu Bulu destroyed the Megamart so that the small population of remaining population of Mimikyu on Akala, the last of their true species.