>>12215127Hariyama could replace Hitmontop, I'm trying to decide between "defending Eagun and the Relic Stone" and "first Shadow Pokémon snagged". I guess he'd be a good parallel with Togekiss.
The Bluno/Verde/Rosso battles don't seem as significant as the others, and which one he battled just opens a whole other can of worms.
Throughout Colosseum, probably, but I'd think he returned Plusle to Duking's care before leaving Orre.