>>45137432Here's the interesting thing; in the Gold, Silver, and Crystal sprites, Smeargle's paint and paw print were brown due to the color limitations, while the shiny sprite had green paint and the entire body had a green hue. The official art made by Sugimori and others, however, always depicted normal Smeargle with green paint. The Gen 3 games reversed the colors so that the normal Smeargle was green and the shiny was a more distinct red to match that Sugimori artwork. This illustration
>>45132961 is somewhere between the two, where the body colors are that of a normal Smeargle but the paint is red. I can't explain that.
>>45137432 is probably the closest answer, since there were Smeargle with other colors of paint in the anime.
