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The Holo members are not forbidden to collab with Holosta and never have been, It's been completely upto the person to decide for at least 2 years now.
The vocal retards who demand Male x Female collabs are the loud obnoxious twitter users who don't watch streams in the first place, Let's not forget that Holo's whole appeal and reason why people liked it in the first place is because they only did girl x girl collabs.
If you want to see male x female collabs, They do plenty of that in Nijisanji.