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What actually happened with Kronii was a little more complicated. Basically she first attempted to understand where her fans were coming from, and then abruptly did an about face and sided with the people calling them incels. It was a betrayal of trust on the deepest level, and whether or not she meant it this way is irrelevant, the effect was the same.
Basically Kronii did not have a lot of things going for her as a streamer early on. She had the Wada model and a nice voice. But she wasn't great at zatsudan, she wasn't great at games, and she wasn't great at singing, she also did karaoke pretty rarely. Which is fine, part of the joy of the Hololive experience is watching your streamer develop over time. Thus people had to find something else to latch onto and they did so by becoming parasocial with her. This wasn't helped by her choosing to watch a lot of "Gosling" movies in member streams like The Truman Show, and of course Ryan Gosling himself in Blade Runner and La La Land.
The male collab thing was not well received by this portion of the fanbase and they made their discontent known through long and lengthy posts in her community, trying to explain themselves as best they could. Whether they were CGDCTchads or just plain unicorns, or people who didn't like the drama that male collabs inevitably bring, they spelled out their grievances, for the most part as composed as they could be. I'd say the only person that came off close to being a unicorn was Aliaga Ayin's post and even that wasn't that deranged. Some of these posts were from long time members, people Kronii would actually talk and react to in chat, Kronies refer to such people as being on the "spreadsheet' which is an alleged list of her "favorite" members. The most notorious of this would be Gabe who was mentally ill and attention seeking from Day 1, but even his post wasn't that crazy.
The point is during a special unarchived member stream, Kronii took the time to read these posts, out loud on stream, in an effort to better understand what these people were unhappy about. And for a brief moment, that won her a lot of love and respect. While Reddit and Twitter were calling these people unicorns and incels, she actually took the time to read them, addressed their concerns, and even defended them, saying she didn't want people attacking her fans. She stated clearly, that while she would NOT stop doing male collabs, she would not initiate them, probably not tweet to them that much, and she would not stream her POV to them. This satisfied all but the most die hard unicorns, and again, even Aliaga Ayin was pacified. The point is, the Kronies that felt this way felt HEARD. She didn't just dismiss them out of hand, like others were doing.
Then less than a day later, she did this tweet, disappeared for a week, and when she came back said she would collab with whoever, whenever, and that was the end of it. There was no further discussion and at the same time the Hololive account made that proclamation against harassing the talents.
So in the eyes of the Kronies who felt betrayed, she had gone from understanding them and defending them, to siding with the people who mocked them and disparaged them, while running and hiding behind the skirts of the company from the inevitable backlash she had to know would follow. Honestly it would have been better if she had never done that unarchived members stream. She gave them hope that she understood, if not fully agreeing with where they came from, and then dashed it violently. That's why they hate her so much. It's not as simple as "male collabs" or "unfortunately idol culture is a thing" it is a real long standing feeling of betrayal and loss from people who watched her for over a year and thought they had a deeper understanding of their oshi and she them.